I forget sometimes that other people that I work with don't use Firefox or Mozilla. I think it's critical for any web developer, but also there are fundamental features that are useful to average users too.
Some of the distinctive features are listed on the Firefox page, but there are two that I'd like to highlight:
My start up page is actually 5 web pages that I need to check on a daily basis. They open in tabs within the one browser window.
Built-in pop-up blocking and image-blocking (blocking images i.e. ads from specific servers).